# timeline of human evolutio contributors to wikimedia project 58–74 minute from wikipedia the free encyclopedi ![[age-of-man-wiki5uz1.jpg|300]] haeckel's paleontological tree of vertebrates (c.1879). the evolutionary history of species has been described as a "tree" with many branches arising from a single trunk. while haeckel's tree is outdated it illustrates clearly the principles that more complex and accurate modern reconstructions can obscur the timeline of human evolution outlines the major events in the evolutionary lineage of the modern human species homo sapiens throughout the history of life beginning some 3.9 billion years ago down to recent evolution within h. sapiens during and since the last glacial perio it includes brief explanations of the various taxonomic ranks in the human lineage. the timeline reflects the mainstream views in modern taxonomy based on the principle of phylogenetic nomenclature; in cases of open questions with no clear consensus the main competing possibilities are briefly outline ## overview of taxonomic rank a tabular overview of the taxonomic ranking of homo sapiens (with age estimates for each rank) is shown belo rank: name: common name: starte life : 3.9 by : archaea3.8 by domain: eukaryota: eukaryotes: 2.1 by : opimoda: excludes plants and their relatives: 1.54 by : amorphea: excludes crums: 1.45 by : obazoa: excludes amoebozoa: 1.3 by : opisthokonta: holozoa + holomycota (cristidicoidea and fungi): 1 by : holozoa: excludes holomycota: 985 my : filozoa: choanozoa + filasterea: 965 my : choanozoa: choanoflagellates + animals: 950 my kingdom: animalia: animals: 665 my subkingdom: eumetazoa: excludes porifera (sponges : parahoxozoa: excludes ctenophora (comb jellies : bilateria: triploblasts / worms: 567 my : nephrozoa: : 558 my : deuterostomia: division from protostomes: 557 my phylum: chordata: chordates (vertebrates and closely related invertebrates): 530 my : olfactores: excludes cephalochordates (lancelets subphylum: vertebrata: fish / vertebrates: 505 my infraphylum: gnathostomata: jawed fish: 460 my : teleostomi: bony fish: 436 my : sarcopterygii: lobe finned fish: 425 my superclass: tetrapoda: tetrapods (animals with four limbs): 395 my : amniota: amniotes (fully terrestrial tetrapods whose eggs are "equipped with an amnion"): 340 my : synapsida: proto-mammals: 318 my : therapsida: limbs beneath the body and other mammalian traits: 280 my class: mammalia: mammals: 220 my subclass: theria: mammals that give birth to live young (i.e. non-egg-laying): 160 my infraclass: eutheria: placental mammals (i.e. non-marsupials): 125 my magnorder: boreoeutheria: supraprimates (most) hoofed mammals (most) carnivorous mammals cetaceans and bats: 115 my superorder: euarchontoglires: supraprimates: primates colugos tree shrews rodents and rabbits: 100 my grandorder: euarchonta: primates colugos and tree shrews: 90 my mirorder: primatomorpha: primates and colugos: 79.6 my order: primates: primates / plesiadapiformes: 66 my suborder: haplorhini: "dry-nosed" (literally "simple-nosed") primates: tarsiers and monkeys (incl. apes): 63 my infraorder: simiiformes: monkeys (incl. apes): 40 my parvorder: catarrhini: "downward-nosed" primates: apes and old-world monkeys: 30 my superfamily: hominoidea: apes: great apes and lesser apes (gibbons): 21 my family: hominidae: great apes: humans chimpanzees gorillas and orangutans - the hominids: 17 my subfamily: homininae: humans chimpanzees and gorillas (the african apes): 12.5 my tribe: hominini: includes both homo and pan (chimpanzees) but not gorilla.: 7 my subtribe: hominina: genus homo and close human relatives and ancestors after splitting from pan - the hominins: 6.1 my (genus): ardipithecus s.l.: : 5.77 my (genus): australopithecus: : 4.3 my genus: homo (h. habilis): : 2.8 my (species): h. erectus s.l.: : 2 my (species): h. heidelbergensis s.l.: : 700000 y species: homo sapiens s.s.: humans: 300000 y date: even 3.9 bya: the earliest life appears possibly as protocells. their genetic material was probably composed of rna capable of both self replication and enzymatic activity; their membranes were composed of lipids. the genes were separate strands translated into proteins and often exchanged between the protocells.further information: abiogenesis rna world earliest known life forms and viral evolutio prokaryotic cells appear; their genetic materials are composed of the more stable dna and they use proteins for various reasons primarily for aiding dna to replicate itself by proteinaceous enzymes (rna now acts as an intermediary in this central dogma of genetic information flow of cellular life); genes are now linked in sequences so all information passes to offsprings. they had cell walls & outer membranes and were probably initially thermophiles.further information: cell (biology) §origin 3.8 bya: last universal common ancestor. all organisms alive today are decended from them. its descendants split into archaea and bacteria 3.55 bya: asgard archaea appea 2.2 bya: origin of the eukaryotes: organisms with nuclei endomembrane systems (including mitochondria) and complex cytoskeletons; they spliced mrna between transcription and translation (splicing also occurs in prokaryotes but it is only of non-coding rnas). the evolution of eukaryotes and possibly sex is thought to be related to the goe as it probably pressured two or three lineages of prokaryotes (including an aerobe one which later became mitochondria) to depend on each other leading to endosymbiosis. early eukaryotes lost their cell walls and outer membranes.further information: eukaryote §origin of eukaryote 1.6 bya: sexual reproduction evolves (mitosis and meiosis) around this time leading to faster evolution where genes are mixed in every generation enabling greater variation for subsequent selectio 1.3 bya: last common ancestor with amoebozo 1 bya: last common ancestor with holomycot 950 mya: the choanozoa lineage of filozoa evolves many features for making cell colonies. last common ancestor with choanoflagellate date: even 665 mya: urmetazoan: the first fossils that might represent animals appear in the 665 million year old rocks of the trezona formation of south australia. these fossils are interpreted as being early sponges. multicellular animals may have existed starting around 800 million years ago 655 mya: separation from the sponge lineage. eumetazoa/diploblast: separation from the ctenophora ("comb jellies") lineage.planulozoa/parahoxozoa: separation from the placozoa and cnidaria lineages. all diploblasts possess epithelia nerves muscles and connective tissue and mouths and except for placozoans have some form of symmetry with their ancestors probably having radial symmetry like that of cnidarians. diploblasts separated their early embryonic cells into two germ layers (ecto- and endoderm). photoreceptive eye-spots evolve 567 mya: ![[xenoturbellajaponica5uz1.jpg|300]] xenoturbella one of the simplest and possibly most primitive living bilateriansurbilaterian: the last common ancestor of xenacoelomorphs protostomes (including the arthropod mollusc and annelid lineages) and the deuterostomes (including vertebrates) (the last two are more related to each other and called nephrozoa). xenacoelomorphs all have a gonopore to expel garmetes but nephrozoans merged it with their anus. earliest development of bilateral symmetry mesoderm head (anterior cephalization) and various gut muscles (and thus peristalsis) and in the nephrozoa nephridia (kidney precursors) coelom (or maybe pseudocoelom) distinct mouth and anus (evolution of through-gut) and possibly even nerve cords and blood vessels. reproductive tissue probably concentrates into a pair of gonads connecting just before the posterior orifice. "cup-eyes" and balance organs evolve (the function of hearing added later as the more complex inner ear evolves in vertebrates). the nephrozoan through-gut had a wider portion in the front called the pharynx. the integument or skin consists of an epithelial layer (epidermis) and a connective layer 558 mya: last common ancestor with protostome 538.8 mya: most known animal phyla appeared in the fossil record as marine species during the cambrian explosion probably caused by long scale oxygenation since around 538.8 million years ago (sometimes called the neoproterozoic oxygenation event or noe) and also an influx of oceanic minerals. deuterostomes the last common ancestor of the chordata lineage hemichordata (acorn worms and graptolites) and echinodermata (starfish sea urchins sea cucumbers etc.) probably had both ventral and dorsal nerve cords like modern acorn worms.an archaic survivor from this stage is the acorn worm sporting an open circulatory system (with less branched blood vessels) with a heart that also functions as a kidney. acorn worms have a plexus concentrated into both dorsal and ventral nerve cords. the dorsal cord reaches into the proboscis and is partially separated from the epidermis in that region. this part of the dorsal nerve cord is often hollow and may well be homologous with the brain of vertebrates. deuterostomes also evolved pharyngeal slits which were probably used for filter feeding like in hemi- and proto-chordate date: even 518 mya: ![[haikouichthys(transparent)5uz1.png|300]] haikouichthys a primitive jawless fish from the cambrian one of the most primitive known vertebratesthe increased amount of oxygen causes many eukaryotes including most animals to become obligate aerobes.the chordata ancestor gave rise to the lancelets (amphioxii) and olfactores. ancestral chordates evolved a post-anal tail notochord and endostyle (precursor of thyroid). the pharyngeal slits (or gills) are now supported by connective tissue and used for filter feeding and possibly breathing. the first of these basal chordates to be discovered by science was pikaia gracilens. other earlier chordate predecessors include myllokunmingia fengjiaoa yunnanozoon lividum and haikouichthys ercaicunensis. they probably lost their ventral nerve cord and evolved a special region of the dorsal one called the brain with glia becoming permanently associated with neurons. they probably evolved the first blood cells (probably early leukocytes indicating advanced innate immunity) which they made around the pharynx and gut. all chordates except tunicates sport an intricate closed circulatory system with highly branched blood vessels.olfactores last common ancestor of tunicates and vertebrates in which olfaction (smell) evolved. since lancelets lack a heart it possibly emerged in this ancestor (previously the blood vessels themselves were contractile) though it could have been lost in lancelets after evolving in early deuterostomes (hemichordates and echinoderms have hearts) ![[sacabambaspiswiki25uz1.png|300]] sacabambaspis an armoured jawless fish (ostracoderm) known from the ordovicianthe first vertebrates ("fish") appear: the agnathans. they were jawless had seven pairs of pharyngeal arches like their descendants today and their endoskeletons were cartilaginous (then only consisting of the chondrocranium/braincase and vertebrae). the jawless cyclostomata diverge at this stage. the connective tissue below the epidermis differentiates into the dermis and hypodermis. they depended on gills for respiration and evolved the unique sense of taste (the remaining sense of the skin now called "touch") endothelia camera eyes and inner ears (capable of hearing and balancing; each consists of a lagena an otolithic organ and two semicircular canals) as well as livers thyroids kidneys and two-chambered hearts (one atrium and one ventricle). they had a tail fin but lacked the paired (pectoral and pelvic) fins of more advanced fish. brain divided into three parts (further division created distinct regions based on function). the pineal gland of the brain penetrates to the level of the skin on the head making it seem like a third eye. they evolved the first erythrocytes and thrombocytes 460-430 ma: ![[reconstructionofentelognathusprimordialisinlateral5uz1.png|300]] life restoration of entelognathus an early placoderm from the silurian epochthe placodermi were the first jawed fishes (gnathostomata); their jaws evolved from the first gill/pharyngeal arch and they largely replaced their endoskeletal cartilage with bone and evolved pectoral and pelvic fins. bones of the first gill arch became the upper and lower jaw while those from the second arch became the hyomandibula ceratohyal and basihyal; this closed two of the seven pairs of gills. the gap between the first and second arches just below the braincase (fused with upper jaw) created a pair of spiracles which opened in the skin and led to the pharynx (water passed through them and left through gills).placoderms had competition with the previous dominant animals the cephalopods and sea scorpions and rose to dominance themselves. a lineage of them probably evolved into the bony and cartilaginous fish after evolving scales teeth (which allowed the transition to full carnivory) stomachs spleens thymuses myelin sheaths hemoglobin and advanced adaptive immunity (the latter two occurred independently in the lampreys and hagfish). jawed fish also have a third lateral semicircular canal and their otoliths are divided between a saccule and utricle 430-410 ma: ![[guiyubw5uz1.jpg|300]] guiyu an early bony fish from the late silurianbony fish split their jaws into several bones and evolve lungs fin bones two pairs of rib bones and opercular bones and diverge into the actinopterygii (with ray fins) and the sarcopterygii (with fleshy lower fins); the latter transitioned from marine to freshwater habitats. jawed fish also possess dorsal and anal fin date: even 390 ma: ![[panderichthysbw5uz1.jpg|300]] panderichthyssome freshwater lobe-finned fish (sarcopterygii) develop limbs and give rise to the tetrapodomorpha. these fish evolved in shallow and swampy freshwater habitats where they evolved large eyes and spiracles.primitive tetrapods ("fishapods") developed from tetrapodomorphs with a two-lobed brain in a flattened skull a wide mouth and a medium snout whose upward-facing eyes show that it was a bottom-dweller and which had already developed adaptations of fins with fleshy bases and bones. (the "living fossil" coelacanth is a related lobe-finned fish without these shallow-water adaptations.) tetrapod fishes used their fins as paddles in shallow-water habitats choked with plants and detritus. the universal tetrapod characteristics of front limbs that bend backward at the elbow and hind limbs that bend forward at the knee can plausibly be traced to early tetrapods living in shallow water.panderichthys is a 90–130cm (35–50in) long fish from the late devonian period (380 mya). it has a large tetrapod-like head. panderichthys exhibits features transitional between lobe-finned fishes and early tetrapods.trackway impressions made by something that resembles ichthyostega's limbs were formed 390 ma in polish marine tidal sediments. this suggests tetrapod evolution is older than the dated fossils of panderichthys through to ichthyostega 375-350 ma: ![[tiktaalikbw5uz1.jpg|300]] tiktaaliktiktaalik is a genus of sarcopterygian (lobe-finned) fishes from the late devonian with many tetrapod-like features. it shows a clear link between panderichthys and acanthostega ![[acanthostegabw5uz1.jpg|300]] acanthosteg ![[ichthyostegabw5uz1.jpg|300]] ichthyostegaacanthostega is an extinct tetrapod among the first animals to have recognizable limbs. it is a candidate for being one of the first vertebrates to be capable of coming onto land. it lacked wrists and was generally poorly adapted for life on land. the limbs could not support the animal's weight. acanthostega had both lungs and gills also indicating it was a link between lobe-finned fish and terrestrial vertebrates. the dorsal pair of ribs form a rib cage to support the lungs while the ventral pair disappears.ichthyostega is another extinct tetrapod. being one of the first animals with only two pairs of limbs (also unique since they end in digits and have bones) ichthyostega is seen as an intermediate between a fish and an amphibian. ichthyostega had limbs but these probably were not used for walking. they may have spent very brief periods out of water and would have used their limbs to paw their way through the mud. they both had more than five digits (eight or seven) at the end of each of their limbs and their bodies were scaleless (except their bellies where they remained as gastralia). many evolutionary changes occurred at this stage: eyelids and tear glands evolved to keep the eyes wet out of water and the eyes became connected to the pharynx for draining the liquid; the hyomandibula (now called columella) shrank into the spiracle which now also connected to the inner ear at one side and the pharynx at another becoming the eustachian tube (columella assisted in hearing); an early eardrum (a patch of connective tissue) evolved on the end of each tube (called the otic notch); and the ceratohyal and basihyal merged into the hyoid. these "fishapods" had more ossified and stronger bones to support themselves on land (especially skull and limb bones). jaw bones fuse together while gill and opercular bones disappear 350-330 ma: ![[pederpes22small5uz1.jpg|300]] pederpespederpes from around 350 ma indicates that the standard number of 5 digits evolved at the early carboniferous when modern tetrapods (or "amphibians") split in two directions (one leading to the extant amphibians and the other to amniotes). at this stage our ancestors evolved vomeronasal organs salivary glands tongues parathyroid glands three-chambered hearts (with two atria and one ventricle) and bladders and completely removed their gills by adulthood. the glottis evolves to prevent food going into the respiratory tract. lungs and thin moist skin allowed them to breathe; water was also needed to give birth to shell-less eggs and for early development. dorsal anal and tail fins all disappeared.lissamphibia (extant amphibians) retain many features of early amphibians but they have only four digits (caecilians have none) 330-300 ma: ![[hylonomusbw5uz1.jpg|300]] hylonomus a primitive relative of amniotes or an early reptile known from the late carboniferousthe late carboniferous saw the emergence of amniota the split between the two major groups of amniotes the ancestors of reptiles and birds (sauropsida) and mammals (synapsida) likely occurred by around 310 million years ago. the skin of the common ancestor of amniotes was covered in reptile-like scales.evolution of the amniotic egg allows the amniotes to reproduce on land and lay shelled eggs on dry land. they did not need to return to water for reproduction nor breathing. this adaptation and the desiccation-resistant scales gave them the capability to inhabit the uplands for the first time albeit making them drink water through their mouths. at this stage adrenal tissue may have concentrated into discrete glands.amniotes have advanced nervous systems with twelve pairs of cranial nerves unlike lower vertebrates. they also evolved true sternums but lost their eardrums and otic notches (hearing only by columella bone conduction ## synapsida and mammali date: even 300-250 ma: ![[palaeohatteriadb5uz1.jpg|300]] palaeohatteria an early sphenacodontian synapsidearly synapsids are referred to as "pelycosaurs" which were dominant land animals during the early permian (300-270 million years ago). they were covered in scaly reptile-like skin. from within the pelycosaurs emerged the therapsids which rose to dominance during the middle and late permian. evidence suggests that early therapsids were covered in pustular hairless skin. the therapsids include the cynodonts the ancestors of mammals which emerged during the late permian. the jaws of cynodonts resembled modern mammal jaws; the anterior portion the dentary held differentiated teeth. this group of animals likely contains a species which is the ancestor of all modern mammals. their temporal fenestrae merged with their orbits. their hindlimbs became erect and their posterior bones of the jaw progressively shrunk to the region of the columella 250-170 ma: cynodonts were diverse and successful animals for much of the triassic period. one subgroup of the cynodonts probainognathia lost the pineal eye and are suggested to have become warm-blooded. advanced probainognathians are likely to have had fur ![[probelesodon5uz1.jpg|300]] chiniquodon a cynodont belonging to probainognathia which includes mammal ![[morganucodon5uz1.jpg|300]] morganucodon an early mammaliaformfrom probainognathia came the first mammals. most early mammals were small shrew-like animals that fed on insects and had transitioned to nocturnality to avoid competition with the dominant archosaurs - this led to the loss of the vision of red and ultraviolet light (ancestral tetrachromacy of vertebrates reduced to dichromacy). although there is no evidence in the fossil record it is likely that these animals had a constant body temperature hair and milk glands for their young (the glands stemmed from the milk line). the neocortex (part of the cerebrum) region of the brain evolves in mammalia at the reduction of the tectum (non-smell senses which were processed here became integrated into neocortex but smell became primary sense). origin of the prostate gland and a pair of holes opening to the columella and nearby shrinking jaw bones; new eardrums stand in front of the columella and eustachian tube. the skin becomes hairy glandular (glands secreting sebum and sweat) and thermoregulatory. teeth fully differentiate into incisors canines premolars and molars; mammals become diphyodont and possess developed diaphragms and males have internal penises. all mammals have four chambered hearts (with two atria and two ventricles) and lack cervical ribs (now mammals only have thoracic ribs).monotremes are an egg-laying group of mammals represented today by the platypus and echidna. recent genome sequencing of the platypus indicates that its sex genes are closer to those of birds than to those of the therian (live birthing) mammals. comparing this to other mammals it can be inferred that the first mammals to gain sexual differentiation through the existence or lack of sry gene (found in the y-chromosome) evolved only in the therians. early mammals and possibly their eucynodontian ancestors had epipubic bones which serve to hold the pouch in modern marsupials (in both sexes) 170-120 ma: ![[juramaiant5uz1.jpg|300]] juramaia sinensis an early therian mammal known from the jurassicevolution of live birth (viviparity) with early therians probably having pouches for keeping their undeveloped young like in modern marsupials. nipples stemmed out of the therian milk lines. the posterior orifice separates into anal and urogenital openings; males possess an external penis.monotremes and therians independently detach the malleus and incus from the dentary (lower jaw) and combine them to the shrunken columella (now called stapes) in the tympanic cavity behind the eardrum (which is connected to the malleus and held by another bone detached from the dentary the tympanic plus ectotympanic) and coil their lagena (cochlea) to advance their hearing with therians further evolving an external pinna and erect forelimbs. female placentalian mammals do not have pouches and epipubic bones but instead have a developed placenta which penetrates the uterus walls (unlike marsupials) allowing a longer gestation; they also have separated urinary and genital openings 100-90 ma: last common ancestor of rodents rabbits ungulates carnivorans bats shrews and humans (base of the clade boreoeutheria; males now have external testicles date: even 90–66 ma: ![[plesiadapis25uz1.jpg|300]] plesiadapi ![[carpolestescl5uz1.png|300]] carpolestes simpsonia group of small nocturnal arboreal insect-eating mammals called euarchonta begins a speciation that will lead to the orders of primates treeshrews and flying lemurs. they reduced the number of mammaries to only two pairs (on the chest). primatomorpha is a subdivision of euarchonta including primates and their ancestral stem-primates plesiadapiformes. an early stem-primate plesiadapis still had claws and eyes on the side of the head making it faster on the ground than in the trees but it began to spend long times on lower branches feeding on fruits and leaves.the plesiadapiformes very likely contain the ancestor species of all primates. they first appeared in the fossil record around 66 million years ago soon after the cretaceous–paleogene extinction event that eliminated about three-quarters of plant and animal species on earth including most dinosaurs.one of the last plesiadapiformes is carpolestes simpsoni having grasping digits but not forward-facing eyes 66-56 ma: primates diverge into suborders strepsirrhini (wet-nosed primates) and haplorrhini (dry-nosed primates). brain expands and cerebrum divides into 4 pairs of lobes. the postorbital bar evolves to separate the orbit from the temporal fossae as sight regains its position as the primary sense; eyes became forward-facing. strepsirrhini contain most prosimians; modern examples include lemurs and lorises. the haplorrhines include the two living groups: prosimian tarsiers and simian monkeys including apes. the haplorrhini metabolism lost the ability to produce vitamin c forcing all descendants to include vitamin c-containing fruit in their diet. early primates only had claws in their second digits; the rest were turned into nails 50-35 ma: ![[aegyptopithecusnt5uz1.jpg|300]] aegyptopithecussimians split into infraorders platyrrhini and catarrhini. they fully transitioned to diurnality and lacked any claw and tapetum lucidum (which evolved many times in various vertebrates). they possibly evolved at least some of the paranasal sinuses and transitioned from estrous cycle to menstrual cycle. the number of mammaries is now reduced to only one thoracic pair. platyrrhines new world monkeys have prehensile tails and males are color blind. the individuals whose descendants would become platyrrhini are conjectured to have migrated to south america either on a raft of vegetation or via a land bridge (the hypothesis now favored). catarrhines mostly stayed in africa as the two continents drifted apart. possible early ancestors of catarrhines include aegyptopithecus and saadanius 35-20 ma: ![[proconsulnt5uz1.jpg|300]] proconsulcatarrhini splits into 2 superfamilies old world monkeys (cercopithecoidea) and apes (hominoidea). human trichromatic color vision had its genetic origins in this period. catarrhines lost the vomeronasal organ (or possibly reduced it to vestigial status).proconsul was an early genus of catarrhine primates. they had a mixture of old world monkey and ape characteristics. proconsul's monkey-like features include thin tooth enamel a light build with a narrow chest and short forelimbs and an arboreal quadrupedal lifestyle. its ape-like features are its lack of a tail ape-like elbows and a slightly larger brain relative to body size.proconsul africanus is a possible ancestor of both great and lesser apes including human date: even 20-15 ma: hominidae (great ape ancestors) speciate from the ancestors of the gibbon (lesser apes) between c. 20 to 16 ma. they largely reduced their ancestral snout and lost the uricase enzyme (present in most organisms) 16-12 ma: homininae ancestors speciate from the ancestors of the orangutan between c. 18 to 14 ma.pierolapithecus catalaunicus is thought to be a common ancestor of humans and the other great apes or at least a species that brings us closer to a common ancestor than any previous fossil discovery. it had the special adaptations for tree climbing as do present-day humans and other great apes: a wide flat rib cage a stiff lower spine flexible wrists and shoulder blades that lie along its back 12 ma: danuvius guggenmosi is the first-discovered late miocene great ape with preserved long bones and greatly elucidates the anatomical structure and locomotion of contemporary apes. it had adaptations for both hanging in trees (suspensory behavior) and walking on two legs (bipedalism) - whereas among present-day hominids humans are better adapted for the latter and the others for the former. danuvius thus had a method of locomotion unlike any previously known ape called "extended limb clambering" walking directly along tree branches as well as using arms for suspending itself. the last common ancestor between humans and other apes possibly had a similar method of locomotion 12-8 ma: the clade currently represented by humans and the genus pan (chimpanzees and bonobos) splits from the ancestors of the gorillas between c. 12 to 8 ma 8-6 ma: ![[sahelanthropuscl5uz1.png|300]] sahelanthropus tchadensishominini: the latest common ancestor of humans and chimpanzees is estimated to have lived between roughly 10 to 5 million years ago. both chimpanzees and humans have a larynx that repositions during the first two years of life to a spot between the pharynx and the lungs indicating that the common ancestors have this feature a precondition for vocalized speech in humans. speciation may have begun shortly after 10 ma but late admixture between the lineages may have taken place until after 5 ma. candidates of hominina or homininae species which lived in this time period include graecopithecus (c. 7 ma) sahelanthropus tchadensis (c. 7 ma) orrorin tugenensis (c. 6 ma) ![[ardi5uz1.jpg|300]] ardipithecusardipithecus is or may be a very early hominin genus (tribe hominini and subtribe hominina). two species are described in the literature: a. ramidus which lived about 4.4 million years ago during the early pliocene and a. kadabba dated to approximately 5.6 million years ago (late miocene). a. ramidus had a small brain measuring between 300 and 350cm3. this is about the same size as the modern bonobo and female chimpanzee brain; it is somewhat smaller than the brain of australopithecines like lucy (400 to 550cm3) and slightly over a fifth the size of the modern homo sapiens brain.ardipithecus was arboreal meaning it lived largely in the forest where it competed with other forest animals for food no doubt including the contemporary ancestor of the chimpanzees. ardipithecus was probably bipedal as evidenced by its bowl shaped pelvis the angle of its foramen magnum and its thinner wrist bones though its feet were still adapted for grasping rather than walking for long distances 4–3.5 ma: ![[bienvenidaalmuseodelucy5uz1.jpg|300]] reconstruction of "lucy"a member of the australopithecus afarensis left human-like footprints on volcanic ash in laetoli northern tanzania providing strong evidence of full-time bipedalism. australopithecus afarensis lived between 3.9 and 2.9 million years ago and is considered one of the earliest hominins - those species that developed and comprised the lineage of homo and homo's closest relatives after the split from the line of the chimpanzees.it is thought that a. afarensis was ancestral to both the genus australopithecus and the genus homo. compared to the modern and extinct great apes a. afarensis had reduced canines and molars although they were still relatively larger than in modern humans. a. afarensis also has a relatively small brain size (380–430cm3) and a prognathic (anterior-projecting) face.australopithecines have been found in savannah environments; they probably developed their diet to include scavenged meat. analyses of australopithecus africanus lower vertebrae suggests that these bones changed in females to support bipedalism even during pregnancy 3.5–3.0 ma: kenyanthropus platyops a possible ancestor of homo emerges from the australopithecus. stone tools are deliberately constructed possibly by kenyanthropus platyops or australopithecus afarensis 3 ma: the bipedal australopithecines (a genus of the subtribe hominina) evolve in the savannas of africa being hunted by megantereon. loss of body hair occurs from 3 to 2 ma in parallel with the development of full bipedalism and slight enlargement of the brai date: even 2.8–2.0 ma: early homo appears in east africa speciating from australopithecine ancestors. the lower paleolithic is defined by the beginning of use of stone tools. australopithecus garhi was using stone tools at about 2.5 ma. homo habilis is the oldest species given the designation homo by leakey et al. in 1964. h. habilis is intermediate between australopithecus afarensis and h. erectus and there have been suggestions to re-classify it within genus australopithecus as australopithecus habilis.ld 350-1 is now considered the earliest known specimen of the genus homo dating to 2.75–2.8 ma found in the ledi-geraru site in the afar region of ethiopia. it is currently unassigned to a species and it is unclear if it represents the ancestor to h. habilis and h. rudolfensis which are estimated to have evolved around 2.4 ma.stone tools found at the shangchen site in china and dated to 2.12 million years ago are considered the earliest known evidence of hominins outside africa surpassing dmanisi hominins found in georgia by 300000 years although whether these hominins were an early species in the genus homo or another hominin species is unknown.further information: homo naledi and homo rudolfensi 1.9–0.8 ma: ![[homo.erectus.adult.female.smithsonian.timevanson.flickr5uz1.jpg|300]] reconstruction of a female h. erectushomo erectus derives from early homo or late australopithecus.homo habilis although significantly different of anatomy and physiology is thought to be the ancestor of homo ergaster or african homo erectus; but it is also known to have coexisted with h. erectus for almost half a million years (until about 1.5 ma). from its earliest appearance at about 1.9 ma h. erectus is distributed in east africa and southwest asia (homo georgicus). h. erectus is the first known species to develop control of fire by about 1.5 ma.h. erectus later migrates throughout eurasia reaching southeast asia by 0.7 ma. it is described in a number of subspecies. early humans were social and initially scavenged before becoming active hunters. the need to communicate and hunt prey efficiently in a new fluctuating environment (where the locations of resources need to be memorized and told) may have driven the expansion of the brain from 2 to 0.8 ma.evolution of dark skin at about 1.2 ma.homo antecessor may be a common ancestor of homo sapiens and neanderthals. at present estimate humans have approximately 20000–25000 genes and share 99% of their dna with the now extinct neanderthal and 95–99% of their dna with their closest living evolutionary relative the chimpanzees. the human variant of the foxp2 gene (linked to the control of speech) has been found to be identical in neanderthals 0.8–0.3 ma: ![[homoheidelbergensis-forensicfacialreconstruction-c5uz1.png|300]] reconstruction of homo heidelbergensisdivergence of neanderthal and denisovan lineages from a common ancestor. homo heidelbergensis (in africa also known as homo rhodesiensis) had long been thought to be a likely candidate for the last common ancestor of the neanderthal and modern human lineages. however genetic evidence from the sima de los huesos fossils published in 2016 seems to suggest that h. heidelbergensis in its entirety should be included in the neanderthal lineage as "pre-neanderthal" or "early neanderthal" while the divergence time between the neanderthal and modern lineages has been pushed back to before the emergence of h. heidelbergensis to about 600000 to 800000 years ago the approximate age of homo antecessor. brain expansion (enlargement) between 0.8 and 0.2 ma may have occurred due to the extinction of most african megafauna (which made humans feed from smaller prey and plants which required greater intelligence due to greater speed of the former and uncertainty about whether the latter were poisonous or not) extreme climate variability after mid-pleistocene transition (which intensified the situation and resulted in frequent migrations) and in general selection for more social life (and intelligence) for greater chance of survival reproductivity and care for mothers. solidified footprints dated to about 350 ka and associated with h. heidelbergensis were found in southern italy in 2003.h. sapiens lost the brow ridges from their hominid ancestors as well as the snout completely though their noses evolve to be protruding (possibly from the time of h. erectus). by 200 ka humans had stopped their brain expansio date: even 300–130 ka: ![[homosapienssapiens(fundortjebelirhoudmarokko)5uz1.jpg|300]] reconstruction of early homo sapiens from jebel irhoud morocco c. 315 000 years bpneanderthals and denisovans emerge from the northern homo heidelbergensis lineage around 500-450 ka while sapients emerge from the southern lineage around 350-300 ka.fossils attributed to h. sapiens along with stone tools dated to approximately 300000 years ago found at jebel irhoud morocco yield the earliest fossil evidence for anatomically modern homo sapiens. modern human presence in east africa (gademotta) at 276 kya. in july 2019 anthropologists reported the discovery of 210000 year old remains of what may possibly have been a h. sapiens in apidima cave peloponnese greece.patrilineal and matrilineal most recent common ancestors (mrcas) of living humans roughly between 200 and 100 kya with some estimates on the patrilineal mrca somewhat higher ranging up to 250 to 500 kya.160000 years ago homo sapiens idaltu in the awash river valley (near present-day herto village ethiopia) practiced excarnation 130–80 ka: marine isotope stage 5 (eemian).modern human presence in southern africa and west africa. appearance of mitochondrial haplogroup (mt-haplogroup) l2 80–50 ka: mis 4 beginning of the upper paleolithic.early evidence for behavioral modernity. appearance of mt-haplogroups m and n. southern dispersal migration out of africa proto-australoid peopling of oceania. archaic admixture from neanderthals in eurasia from denisovans in oceania with trace amounts in eastern eurasia and from an unspecified african lineage of archaic humans in sub-saharan africa as well as an interbred species of neanderthals and denisovans in asia and oceania 50–25 ka: ![[homosapienssapiensoaserumänien(danielahitzeman5uz1.jpg|300]] reconstruction of oase 2 (c. 40 ka)behavioral modernity develops by this time or earlier according to the "great leap forward" theory. extinction of homo floresiensis. m168 mutation (carried by all non-african males). appearance of mt-haplogroups u and k. peopling of europe peopling of the north asian mammoth steppe. paleolithic art. extinction of neanderthals and other archaic human variants (with possible survival of hybrid populations in asia and africa). appearance of y-haplogroup r2; mt-haplogroups j and x after 25 ka: ![[homosapiens-neolithic-reconstruction-muse5uz1.jpg|300]] reconstruction of a neolithic farmer from europe science museum in trentolast glacial maximum; epipaleolithic / mesolithic / holocene. peopling of the americas. appearance of: y-haplogroup r1a; mt-haplogroups v and t. various recent divergence associated with environmental pressures e.g. light skin in europeans and east asians (kitlg asip) after 30 ka; inuit adaptation to high-fat diet and cold climate 20 ka.extinction of late surviving archaic humans at the beginning of the holocene (12 ka). accelerated divergence due to selection pressures in populations participating in the neolithic revolution after 12 ka e.g. east asian types of adh1b associated with rice domestication or lactase persistence. the past 100000 years have seen selective reductions in brain size for some human lineages during warmer interglacial period **+** evolution of human intelligenc **+** human evolutio **+** recent human evolutio **+** genetic history of pleistocene human **+** list of human evolution fossil **+** human histor **+** natural histor **+** history of eart **+** timeline of prehistor **+** timeline of the evolutionary history of lif **+** chronology of the univers **+** list of timeline 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